Checkpoints

A bit long sentence, so please be patient.
Thank you.

I think:

"The increased recognition of the role of B cells in autoimmunity has led to renewed interest in the checkpoints that govern B cell tolerance, identifiable as ‘central’ when established in the bone marrow and ‘peripheral’ after B cells have exited to the periphery."

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The increased recognition of the role of B cells in autoimmunity has led to renewed interest in the checkpoints that govern B cell tolerance, identifiable as ‘central checkpoints’ when B cells were established (or "when B cells appear") in the bone marrow and ‘peripheral checkpoints’ after B cells have exited to the periphery.

Am I on the right track?

Context:

The increased recognition of the role of B cells in autoimmunity has led to renewed interest in the checkpoints that govern B cell tolerance, identifiable as ‘central’ when established in the bone marrow and ‘peripheral’ after B cells have exited to the periphery. Genetically determined defects that have an impact in either site can result in malfunctioning B cells and/or the accumulation of autoreactive B cells in the blood (Eric Meffre Yale University, New Haven, CT).