@wandeljw,
Thats the point of the work by Venditti. He and his team merely computerized time elements before and after major "speciation pulses" and gelogical events .
Years ago I posted a thread that was entitled "Is evolution adaptive"?. According to many scientists the answer was clearly yes but the mechanism of natural selection was "in the way" to a purely mechanistic evaluation. Its interesting to see how the entire topic of evolutionary synthesis will be modified.
As I was never a proponent of Punctuated Equilibrium since (IMHO) it was merely an artifact of an incomplete geological record, now maybe Im gonna have to modify a bit since adaptive speciation seems to be the major driver.
In many cases we see speciation occur with a modification that resulted from one bug genetic change (which recorded a successful adaptation and this was passed on). An example of that is
achondroplasia which is a single jump mutation that results in animals with shortened legs in one swoop. We know of this genetic expression from natuarl speciation as well as in artificial selection to make short legged dogs like daschunds and bassett hounds. This same thing occured (probably) when the environment around present day PAkistan led to the inundation of the area and the pre-cetacean ancestors became suddenly modified to prefer shorter legs which , together with changes in tne toe structures , led to a long legged hooved animal quickly leaving fossils of short legged many toed animals like pachycetus.
GEOLOGICAL ENVIRONMENT RULES!!